Ascosporogenesis in Tuber magnatum is intermediate between the general patterns of Euascomycetes and that of Hemiascomycetes. T. magnatum ascospores are delimited individually by two membranes which originate from the ascus plasmalemma; wall material is deposited between these membranes. The epispore is deposited first. The outer perispore is initially expanded but then contracts and fills with increasingly compact electron-dense material to form the surface ornamentations. The electron-transparent endospore is deposited inside the epispore.
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